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Ubstitution of Lys-1149, to the Win1 MAPKKK to test whether Win1 plays a more important role in osmostress signaling than Wis4 (Samejima et al., 1997, 1998). The strain expressing the catalytically inactive Win1KM showed Spc1 activation equal to that in the wis4KM strain, and only the wis4KM win1KM double mutant failed to induce activation of Spc1 after osmostress (Figure 3B). The KM mutations in
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